About the Journal

The Philippine Journal of Systematic Biology is the official publication of the Association of Systematic Biologists of the Philippines. It was founded during the Annual Conference of the Association of Systematic Biologists in 2007 held at the National Museum of the Philippines. It is the spiritual successor of the ASBP Communications that was published from 1984-1992.
SCImago Journal & Country Rank
April 30, 2019

The PJSB has been accepted for indexing by the ASEAN Citation Index. ASEAN Citation Index (ACI) is a central regional database which was designed and set up to index all the bibliographic records and the citations of all quality ASEAN research outputs appeared in the ASEAN scholarly journals (https://www.asean-cites.org/index.php?r=home).

March, 2019

To all accepted Oral and Poster Presentors of the 2nd PSFBE:

You are invited to submit full papers for consideration in a special issue of the Philippine Journal of Systematic Biology

All submitted papers will be evaluated by the editors of the PJSB and will be subjected to peer-review prior to acceptance. Deadline is on June 30, 2019.

News Archive
May 5, 2017

As per Commision on Higher Education CMO 50 series of 2017, the Philippine Journal of Systematic Biology is now a CHED Recognized Journal.

September 10, 2017

The PJSB is now a member of the Committee on Publication Ethics (COPE). COPE was founded in 1997 by a group of medical journal editors concerned about publication misconduct, e.g. plagiarism, attempted or actual redundant publication, attempts to pass off
fraudulent data, unethical research, breaches of confdentiality, etc. Originally a loose gathering of individuals COPE is now a limited company and registered charity in the UK.

July 5, 2018

The PJSB is now indexed in Scopus.  Scopus is one of the largest indexing database of peer-reviewed publications managed by Elsevier.

Focus and Scope
The Philippine Journal of Systematic Biology  (Print ISSN: 1908-6865; Online ISSN: 2508-0342)  is an annual, peer-reviewed journal in English that publishes high quality reports of original research and reviews in the field of Taxonomy & Systematics, Ecology and Conservation Biology. The PJSB is a CHED recognized journal under CHED CMO 50 series of 2017. It is also included in the Clavariate Analysis (formerly a subsidiary of Thomson Reuters) Master Journal List under Zoological Record, BIOSIS Previews and Biological Abstracts.The PJSB is the official publication of the Association of Systematic Biologists of the Philippines (SEC Registration: Association of Philippine Taxonomists, Inc.). The organization financially supports the journal and all its endeavors. The journal aims to build up quality information on animal, plant and microbial diversity in the Philippines. Articles dealing with original research or reviews in Zoological, Botanical or Microbial Systematics, Ecology and Conservation Biology are welcome.  Authors are advised to consult a recent issue of PJSB for the current format and style.  All manuscripts are reviewed by at least two qualified referees.

The PJSB strongly adheres to research and publication ethics. Submitted manuscripts should not have been published elsewhere nor should have been submitted for review / publication simultaneous with the submission in PJSB.

Peer Review Process
• Preliminary evaluation on the scope and style of the manuscript will be done by two handling editors expert in the field.
• Papers that adhere to the journal scope and style will be sent to 2-3 peer reviewers suggested by two handling editors through the managing editor. The PJSB follows a “double-blind” review on all submitted manuscripts, withholding the names of the authors as well as the identity of the reviewers.
• Manuscripts returned to the authors are given 2-3 weeks to be revised. Upon return of the manuscript, the revised paper should be accompanied by a letter highlighting the revisions or answers to the reviewers queries.
• Final decision on the acceptance or rejection of papers are made by the two handling editors.
Open Access Policy
This journal provides immediate open access to its content on the principle that making research freely available to the public supports a greater global exchange of knowledge. PJSB articles and other forms of publications published by the journal is protected under the Intellectual Property Code of the Philippines. Articles are distributed under the terms of the Creative Commons Attribution Non Commercial License (http://creativecommons.org/licenses/by-nc/3.0/), which permits unrestricted non-commercial use, distribution, and reproduction in any medium, provided the original work is properly cited.
Author Guidelines

General Manuscript Format
• Manuscripts must be submitted as an MS Word file (.doc) attachment to the Managing Editor at pjsb2016@yahoo.com. Submissions should include a cover letter explicitly stating that the manuscript has not been submitted elsewhere for review or has not yet been published.
• The manuscript must be double-spaced using 12 pts, Times New Roman with a margin of 2.5 cm (1 in.) on all sides with row numbers. Number all pages consecutively on the top right corner, including those carrying tables, figures, legends and figures. All measurements should be given in metric unit.

• The manuscript must be written in English following Standard English grammar. Authors are encouraged to have their manuscript checked for grammatical errors by an English editor before submission.

Manuscript Sections Format
• Title Page
o Includes the title (ALL CAPITAL LETTERS, BOLDFACE), full name of each author (without academic degrees) (ALL CAPITAL LETTERS) and up to five keywords (italicized) without repeating those already in the title. Indicate the corresponding author with an asterisk and indicate his email address after the institutional affiliation.

• Abstract
o This occupies page 2 of the manuscript. Each paper should be provided with a summary / abstract not exceeding 300 words and written in one paragraph. It is suggested that the abstract should start with the research targets, followed by the main methods used, major results and the implications of the findings that may be of interest to the scientific community. Please provide 4 to 6 keywords. Keywords should not include any term that already appears in the title.

• Text
The main body of the manuscript starts on the 3rd page. Major headings should be typed in ALL CAPITAL LETTERS, BOLDFACE while subheadings are boldface and italicized. For Review Papers, the summary is followed by a short introduction, the review proper (with subheadings) and the Literature Cited. Full Research papers includes the following major sections:
 Introduction. A short discussion on the objectives of study.
 Materials and Methods. Experimental procedure.
 Results and Discussion. Results should be accompanied by tables, graphs, pictures.
 Taxonomy / Systematics. For papers dealing with descriptions / re descriptions of new taxa
 Conclusions and Recommendations. Conclusions derived from the results and possible future modifications the authors suggest.
 Acknowledgements should be brief and should primarily indicate financial support and / personal assistance in the work being published. Do not acknowledge anonymous reviewers.
 Literature cited. References used and cited in the study. These should be listed alphabetically. Avoid citing gray literature or reference not accessible through indexes / libraries. Write journal / book titles in full. It is the author’s responsibility to check that all papers cited in the manuscript appear in the Literature Cited section.

Article in Journal
Adorador, V.C., M.A. Bernardo, J.L.L. Gonzales, A.C. Lam, F.A.M. Malijan, R.D.S. Papa & J.R. Castillo, 2006. Morphological and DNA size-based differences among Corallana spp. in Oreochromis sp. found in open waters and fish cages in Taal Lake. The Philippine Biota, 39: 5-11.

Book Section
Zafaralla, M.T., C.R. Barril, A.C. Santos-Borja, E.V. Manalili, J.T. Dizon, J.G. Sta. Ana & N.O. Aguilar, 2005. Water Resources. In: Lasco, R.D. & M.A.O. Espaldon, (ed.), Ecosystems and People: The Philippine Millennium Ecosystem Assessment (MA) Sub-global Assessment. Environmental Forestry Programme, College of Forestry and Natural Resources, University of the Philippines Los Baños, Los Baños, Laguna. pp. 65- 112.

Fernando, C.H., 2002. A Guide to Tropical Freshwater Zooplankton -Identification, Ecology and Impact on Fisheries. Backhuys Publishers, Leiden, the Netherlands. 291 pp.

Notes on Figures and Tables

*Tables and figures must be submitted separately from the word file. Only table and figure labels must appear after the references
Tables. Tables must be submitted as separate word files. All tables should be prepared with tabulators (without preceding/trailing spaces). If you prefer using the table tool of your word-processing programme, do not insert line breaks (¶) and/or preceding/trailing spaces inside cells (i.e. there can be only one line of text within a cell). New table rows should be created by adding new rows of table cells (even if some cells are left empty), not by hitting enter and creating a new lines within existing cells. CR symbols (¶) cannot be preset in any table cell. The number of cells in a table should equal the number of rows multiplied by the number of columns. This must be done on a separate word file from the main manuscript. Do not use vertical lines to separate columns. Avoid crowded tables. Do not present data in both graphical and tabular form. Tables should be self-explanatory / understandable without reference to text. Table headings should be placed on top of the table. See Example

Figures. Line drawings, computer plots or photographs should be submitted separately as digital files in jpeg, or tiff format. The images should clearly show intended details to support the manuscript. Picture quality must not be below 5 megapixels/640X480 resolution/200kb/1200 dpi (for scanned images). File names should indicate the following: surname of primary author, year of submission, the word Fig followed by the number (ex. Perez2009Fig1.jpeg). Provide figure captions in a separate sheet at the end of the manuscript. Figure plates/panels should be labeled (a), (b), (c), etc. and referred to in the text as, for example, Fig. 1a.

Ethical responsibilities of Authors
• The PJSB is a member of  the Committee on Publication Ethics and practices high ethical standards on submittted manuscripts and review process. Papers submitted to the PJSB must not yet be submitted or published to other journals and all co-authors and their affiliations have approved its submission and publication. The PJSB will not be held legally responsible should there be any claims for compensations and the like.
• The paper must not be a single study that are split up into several parts to increase the quantity of submission to various journals or to the PJSB
• No data including images have been fabricated or manipulated to support the authors conclusions
• Proper acknowledgement of other researches should be done by the authors in any part of their manuscript. Any form of plagiarism is not tolerated by the PJSB
• All authors have contributed sufficiently to the submitted paper thus will collectively share responsibility and accountability for the results.
• Changes in authorship or order of the authors will not be accepted after the acceptance for publication.
• Change in authorship in any form must be justified with a letter addressed to the Editor-in-Chief.
• The authors should be prepared to send relevant documents or data upon request by the PJSB to verify the validity of the results.
• Any breach of any of the ethical procedures stated here will result into the immediate rejection of the manuscript and legal action will be taken by the PJSB if necessary. The PJSB adopts the Committee on Publication Ethics(COPE) rules and guidelines in both the peer review process and submission of manuscripts.
Disclosure of Potential Conflicts of interest
• Authors must declare all relations or interests that may have possible influence or impart bias on their publications. The PJSB respects the right of the authors and will not force them to release information if they feel there is no conflicts of interest in the publication of their work. Interests that go beyond financial interests or other forms of compensation that is deemed important to the readers should be disclosed by the authors as well.
• Potential conflicts of interest may be directly or indirectly related to their published work may include but are not limited to the following:
o Research Grants (indicate funding agency and grant number in the acknowledgement section)
o Financial award or support received from educational programs, presenting the research in a symposia or affiliation
o Employment
o Support from a project sponsor
o Position on advisory board or board of directors or other type of management relationships
o Multiple affiliations
o Financial Relationships, such as investment interests
o Intellectual property rights
o Filial, Familial or any other relation that may have financial interest in the work/publication
*If there are any Conflicts of interest in the study the authors must include a “Disclosure of conflicts of interest” after the Acknowledgement section or before the Reference Section.
Section Policies
Manuscripts submitted to the PJSB are categorized according to content and scientific contributions. PJSB classifies submitted manuscript into four categories:
o Full Research Papers
o Review Papers, Taxonomic Revisions, Opinion Papers
o Short Notes/Communications
o Book Reviews
Publication Frequency
• The Philippine Journal of Systematic Biology is published annually. PDF’s are posted upon finalization of layout within the year of intended publication.
Editorial Board
Lawrence Liao, Ph.D. (Hiroshima University, Japan)


Domingo Madulid, Ph.D. (De La Salle University-Manila, Philippines)

Editorial Board
Domingo Madulid, Ph.D. (De La Salle University-Manila, Philippines)
Lawrence Liao, Ph.D. (Hiroshima University, Japan)
Hidetoshi Nagamasu, PhD (Kyoto University Museum, Japan)
Boon-Chuan Ho, Ph.D. (Singapore Botanic Gardens,Singapore)
Monica Suleiman, Ph.D. (Universiti Malaysia Sabah, Malaysia)

Molecular Phylogenetics/Evolutionary Biology
Grecebio Jonathan D. Alejandro, Ph.D. (University of Santo Tomas, Philippines)
Irene Samonte Padilla, Ph.D. (Max-Planck-Institut für Evolutions biologie, Germany)
Mike Thiv, Ph.D. (Staatliches Museum fuer Naturkunde, Germany)
Ga Hun Boo, Ph.D. (Chungnam National University, South Korea)

Irineo Dogma, Jr., Ph.D. (University of Santo Tomas, Philippines)
Windell Rivera, Ph.D. (University of the Philippines – Diliman, Philippines)
Yuan Kun Lee, Ph.D. (National University of Singapore, Singapore)

Ornithology/Mammalogy/Herpetology/Invertebrates/Conservation Biology
Juan Carlos Gonzalez, Ph.D. (University of the Philippines – Los Baños, Philippines)
Suchai Worachananant, Ph.D. (Kasetsart University, Thailand)
Eric Rickart, Ph.D. (University of Utah, United States of America)
Rafe Brown, Ph.D. (University of Kansas, United States of America)
Terrence Gosliner, Ph.D. (California Academy of Sciences, United States of America)
Julia T. Sigwart, Ph.D. (Queen’s University, United Kingdom)
Tin-Yam Chan, Ph.D. (National Taiwan Ocean University, Taiwan)

Aimee Lyn Barrion-Dupo, Ph.D.  (University of the Philippines – Los Baños, Philippines)
Anindo Choudhury, Ph.D. (St. Norbert’s College, United States of America)
Masahiro Ohara, Ph.D. (Hokkaido University Museum, Japan)

Hydrobiology/Icthyology/Zooplankton Ecology and Systematics/Limnology
Roberto Pagulayan, Ph.D. (Angeles University Foundation, Philippines)
Rey Donne Papa, Ph.D. (University of Santo Tomas, Philippines)
Hiroyuki Motumura Ph.D. (Kagoshima University Museum, Japan)
Jeffrey Kwik, Ph.D. (National University of Singapore, Singapore)
Henri Dumont, Ph.D. (Jinan University, China)
Bo-Ping Han, Ph.D. (Jinan University, China)
Maria Holynska, Ph.D. (Polish Academy of Sciences, Poland)

Editorial Team

Eric Zeus C. Rizo, Ph.D. (University of the Philippines-Visayas, Philippines) – Managing Editor
Jayson G. Chavez, Ph.D. (Far Eastern University, Philippines) – Assistant Managing Editor
Maria Isabella J. Escobar, M.Sc. (University of Santo Tomas, Philippines) – Lay-out Editor

Compliance with Ethical standards
• To promote objectivity, transparency and compliance to accepted ethical principles and professional conduct, the authors should include an appendix on the relevant information regarding, conflict of interest whether financial or not, government permits on the collection of wildlife, and a statement on the welfare of animals if the research involved experimentation on animals. Information on funding sources must be clearly indicated in the acknowledgement section.
• The corresponding author is responsible for gathering documentation of compliance with ethical standards upon request during the peer review process or after publication
• The PJSB reserves the right to reject manuscript that do not comply to stated ethical procedures and responsibilities. The authors will be held responsible for any falsification of information related to compliance on ethical standards.

• Complaints on the peer-review process, plagiarism, falsification and fabrication of data and the like, conflicts of interests and other publication and research malpractice must be sent to the Managing Editor of the PJSB (pjsb2016@yahoo.com) in a letter addressed to the Editor-in-Chief.
• If the complaint proves to have merit, the Editorial Board will process the complaint and will act on it accordingly. If allegations on research misconduct is proven, manuscript maybe rejected, given an addendum upon publication of manuscript (if it has been accepted), banning of authors to publish in the PJSB or other legal action that the PJSB need to protect its integrity and adherence to standard ethical publication and research practices and procedures.

Charges on Publication for Non ASBP Members
• PJSB has no page charges for articles with at least one bona fide member of the ASBP as author. Papers without ASBP author-members are still encouraged to submit but will be charged a corresponding minimal publication fee upon acceptance of the manuscript.
o Non-ASBP Member local- Php 3000.00
o Non-ASBP Member foreign – USD 100.00

ASBP Communications  Archive

Philippine Journal of Systematic Biology

Volume 14 Issue 3

Table of ContentsButterfly range extensions in the Philippines and Indonesiaby Jade Aster T. Badon and David J. LohmanPhilippine Journal of Systematic Biology, Volume 14, Issue 3, 2020, DOI: 10.26757/pjsb2020c14001Date Posted (Final Published Version) :August 12, 2020...

Volume 14 Issue 2-Special Issue No.2

Table of ContentsEchoes from the past – the identity of Sundathelphusa picta (von Martens, 1868) and descriptions of two new species of freshwater crab (Crustacea, Brachyura, Gecarcinucidae, Sundathelphusa) from Luzon Island, the Philippinesby Peter K. L. Ng and Jose...

Volume 13 Issue 1 (2019)

Table of ContentsNew records of Rosenvingea (Scytosiphonaceae, Phaeophyceae) from the Philippinesby Wilfred John E. Santiañez and John A. WestPhilippine Journal of Systematic Biology, Volume 13, Issue 1, 2019, Pages 1-8, DOI:10.26757/pjsb2019a13001Date Posted (Final...

Volume 12 Issue 2 (2018)

Table of ContentsHistorical trends among cestode research of vertebrates in the Philippines by Vanessa V. Martinez and Jonathan Carlo A. Briones Philippine Journal of Systematic Biology, Volume 12, Issue 2, 2018, Pages 1-12Date Posted (Final Published Version) :...

A Special Tribute to Dr. Benito Tan (Volume 12 Issue 1) (2018)

Guest Editor: Lawrence Liao Ph.D. Table of Contents A tribute to Benito C. Tan (1946–2016) Boon-Chuan Ho & James R. Shevock Rosario R. Rubite Elena Mencias-Ragragio Jef Mancera Download PDF Amphi-Pacific tropical disjunctions in the bryophyte floras of Asia and...

Volume 11 Issue 2 (2017)

Table of Contents Comparative Pollen Viability and Pollen Tube Growth of two Endemic Philippine Etlingera (Zingiberaceae, Alpinioideae) by Noe P. Mendez, Heidi C. Porquis, Evangeline B. Sinamban, and Florfe M. Acma Philippine Journal of Systematic Biology, Volume 11,...

Volume 10 (2016)

Table of Contents Rafflesia camarinensis (Rafflesiaceae), A New Species from Mt. Asog, Camarines Sur, Luzon Island, Philippinesby Flordeliz B. Valenzuela, Gertrudes Girlie Jaucian-Adan, Esperanza Maribel G. Agoo and Domingo A. MadulidPhilippine Journal of Systematic...

Volume 9 (2015)

Table of Contents A critical review of the taxonomic status of Rafflesia philippensis Blanco (Rafflesiaceae) from the Philippines by Domingo A. Madulid and Esperanza Maribel G. Agoo Philippine Journal of Systematic Biology, Volume 9, 2015, Pages 1-9 Abstract An...

Volume 8 (2014)

Table of Contents A new cave-dwelling mygalomorph spider of the genus Phlogiellus Pocock, 1897 (Araneae: Theraphosidae: Selenocosmiinae) from Burdeos, Polillo Island, Quezon Province, Philippinesby Aimee Lynn A. Barrion-Dupo, Alberto T. Barrion and Joseph B....

Volume 7 (2013)

Table of Contents A new record of Pyrostria (Vanguerieae-Rubiaceae) from the Philippines inferred from molecular and morphological databy Grecebio Jonathan D. Alejandro, Elizabeth H. Arenas, Ma. Chiela M. Cremen and Axel H. ArriolaPhilippine Journal of Systematic...

Volume 6 (2012)

Table of Contents Morphological and molecular identification of a novel species of Uvaria (Annonaceae) with potential medicinal propertiesby P.G.D. Cabuang, B.S. Exconde, V.I.M. Lim, D.K.M. Padilla, S.R. Salas, A.P.G. Macabeo, B.O.C. Lemana, and   G.J.D....

Volume 5 (2011)

Table of Contents Methylobacterium zatmanii, a pink pigmented facultative methylotrophic (PPFM) bacterium isolated from the human oral cavity by Thaddeus M. Carvajal, Ronabelle L. Tan And Anthony C. Lee Philippine Journal of Systematic Biology, Volume 5, 2011, Pages...

Volume 4 (2010)

Table of Contents Characterization of marine yeasts isolated from different substrates collected In Calatagan, Batangasby Carly Simon P. Ramirez, Connel O. Go, Sophia Angelica S. Hernandez, Hanah I. Ruiz, Maureen B. Sabit and          Thomas Edison E. Dela Cruz...

Volume 3 (2009)

Table of Contents An enumeration of the woody plants of Cantipla forest fragments, Cebu Island, Philippines by Geofe O. Cadiz & Inocencio E. Buot, Jr. Philippine Journal of Systematic Biology, Volume 3, 2009, Pages 1-7 Abstract Woody plants in Cantipla forest...

Volume 2 (2008)

Table of Contents Tree elements along the western slope of Mt. Lobo: species composition from 700 to 1, 007m altitudes by Anacleto M. Caringal Philippine Journal of Systematic Biology, Volume 2, 2008, Pages 4-11 Abstract Mt. Lobo, the highest peak in the coastal...

Volume 1 (2007)

Table of Contents Pteridophyte and gymnosperm diversity in Musuan, Bukidnon by Victor AmorosoPhilippine Journal of Systematic Biology, Volume 1, 2007, Pages 1-14Abstract Inventory of pteridophytes and gymnosperms in Musuan, Bukidnon revealed 102 species, 52 genera and...